Affiliations

• Research Professor, Life Sciences Institute
• G. Robert Greenberg Distinguished University Professor, Department of Biological Chemistry, UM Medical School
• Professor, Department of Chemistry, College of Literature, Science & the Arts
• Interdepartmental Professor & Program Committee Member, Program in Medicinal Chemistry, UM College of Pharmacy
• Search Committee Member, Biological Sciences Scholars Program
• Faculty Mentor, Molecular Biophysics Graduate Training Program
• Faculty Mentor, Cellular Biotechnology Graduate Training Program
• Faculty Mentor - Biological Chemistry, Pharmocological Sciences Training Program
• Faculty Mentor, Medical Scientist Training Program, UM Medical School

Education

• BA, Biology (Summa Cum Laude), Radcliffe College, 1960
• PhD, Biophysics, University of Michigan, 1969

Honors and Awards

• 1994: Distinguished Biomedical Faculty Lectureship, University of Michigan
• 1997: Distinguished Faculty Achievement Award, University of Michigan
• 2000: American Society for Biochemistry & Molecular Biology William A. Rose Award
• 2001: American Chemical Society, Biological Chemistry Division, Repligen Award
• 2001: 12th International Symposium on Chemistry and Biology of Pteridines Gowland Hopkins Award
• 2003: Henry Russel Lecturer, University of Michigan

Memberships and Offices in Professional Societies

• 1959- : Phi Beta Kappa
• 1959- : American Society for Biochemistry & Molecular Biology
• 1997-2001: Michigan Society of Fellows (Senior Fellow)
• 1998- : American Association for the Advancement of Science (Fellow)
• 2002- : American Society of Microbiology (Fellow)
• 2002- : National Academy of Sciences
• 2004- : Institute of Medicine